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What is the "@" in my wl-summary-line?

To: Wanderlust Mailing List <wl-en@lists.airs.net>
Subject: What is the "@" in my wl-summary-line?
From: David Abrahams <dave@boostpro.com>
Date: Wed, 13 Oct 2010 11:53:00 -0400
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I am seeing "[@]" in my summary line occasionally, and I don't know
what it means.  My wl-summary-line-format is set to
"%T%P%M/%D(%W)%h:%m %[ %17f %]%[%1@%] %t%C%s"

Can anyone explain this for me?
Thanks!

-- 
Dave Abrahams
BoostPro Computing
http://www.boostpro.com

Follow-Ups:
- Re: What is the "@" in my wl-summary-line?
  - From: Erik Hetzner <ehetzner@gmail.com>

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